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Copy of Endogenous Group Formation experiment

Outline of experiment to be given to groups in USA
by

Stephen Heap

on 1 March 2016

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Transcript of Copy of Endogenous Group Formation experiment

How does
developmental
history influence individual behaviour?
How does childhood
environment
affect
proximate
responses to social interactions?
What
ultimate
explanations could apply?
Mikael Puurtinen
Stephen Heap
Jaakko Junikka
How do
individual behaviours
and
group level processes
interact to affect selection at a
genetic level
?
Punishment
shifts selection
from individual to
group level
Group competition
increases
and
stabilises
within group cooperation

How rational agents perform
How humans performed
Made
costly
in the third experiment
Group size
Cooperation
More incentive for selfishness
Group strength in competition
Bigger groups generate more public goods
Group size
Group size
Individual returns from group competition
Individuals get less spoils in bigger groups
But group competition may keep cooperation in check
May be
critical group sizes
at which cooperation breaks down or groups collapse
Arms races
Selection for smaller groups
Selection for larger groups
Selfish individuals limit their expected fitness by
harming
their group
Cooperation
breaks down
in isolated groups
Punishment is
paradoxically
common even when it can be
costly
to individuals and groups
Cooperative
groups have selective
advantages
over non-cooperative groups
Cooperation
is an individual trait that is selected for due to
conflict
between
groups
Cooperation and conflict are the keys

for understanding
organisation and complexity in biology

Dependence on group performance
Fitness
Selfish strategy
Cooperative strategy
Individual fitness =
vs
+
Fitness from within-group competition
vs
}
Fitness from between-group competition
vs
+
vs
Selfish
strategy pays
Cooperative
strategy pays
}
=
Individual fitness =
vs
vs
+
}
}
}
}
}
}
Cooperation
: individual action that improves the fitness of another
There is no
cooperation
without
conflict

at a higher organisational level

Costly punishment
prevails
in group competition and
equalises
earnings
within
groups

Group competition
emerges
spontaneously in randomly formed groups and
maintains
a stable level of cooperation

How does
group competition
affect the relationship between
cooperation
and
population structuring
?

Complex population structures
can be determined by
individual level decisions
At the individual level we have...
...
cooperative
behaviour can evolve...
... which influence whether or not...
... behaviours that
structure
the population...
... which then affects the payoffs of...
1. The size of group that they want to join
2. How much to invest in the public good
3. Vote on how to interact with other groups
Full transcript